AT4G37180 cDNA ORF clone, Arabidopsis thaliana(thale cress)

The following AT4G37180 gene cDNA ORF clone sequences were retrieved from the NCBI Reference Sequence Database (RefSeq). These sequences represent the protein coding region of the AT4G37180 cDNA ORF which is encoded by the open reading frame (ORF) sequence. ORF sequences can be delivered in our standard vector, pcDNA3.1+/C-(K)DYK or the vector of your choice as an expression/transfection-ready ORF clone. Not the clone you want? Click here to find your clone.

***CloneID Accession No. Definition **Vector *Turnaround time Price (USD) Select
OAb17315 NM_179182.2
Latest version!
Arabidopsis thaliana Homeodomain-like superfamily protein (AT4G37180), mRNA. pcDNA3.1-C-(k)DYK or customized vector 7-9 $258.30
$369.00
OAb17316 NM_119881.3
Latest version!
Arabidopsis thaliana Homeodomain-like superfamily protein (AT4G37180), mRNA. pcDNA3.1-C-(k)DYK or customized vector 7-9 $258.30
$369.00
OAb40114 NM_001342435.1
Latest version!
Arabidopsis thaliana Homeodomain-like superfamily protein (AT4G37180), mRNA. pcDNA3.1-C-(k)DYK or customized vector 9-11 $174.30
$249.00

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** You may select a custom vector to replace pcDNA3.1+/C-(K)DYK after clone is added to cart.

** GenScript guarantees 100% sequence accuracy of all synthetic DNA constructs we deliver, but we do not guarantee protein expression in your experimental system. Protein expression is influenced by many factors that may vary between experiments or laboratories. In addition, please pay attention to the signal peptide, propeptide and transit peptide in target ORF, which may affect the choice of vector (N/C terminal tag vector).

***One clone ID might be correlated to multiple accession numbers, which share the same CDS sequence.

  • Reference Sequences (Refseq)
    CloneID OAb17315
    Clone ID Related Accession (Same CDS sequence) NM_179182.2
    Accession Version NM_179182.2 Latest version! Documents for ORF clone product in default vector
    Sequence Information ORF Nucleotide Sequence (Length: 1092bp)
    Protein sequence
    SNP
    Vector pcDNA3.1-C-(k)DYK or customized vector User Manual
    Clone information Clone Map MSDS
    Tag on pcDNA3.1+/C-(K)DYK C terminal DYKDDDDK tags
    ORF Insert Method CloneEZ™ Seamless cloning technology
    Insert Structure linear
    Update Date 2019-02-13
    Organism Arabidopsis thaliana(thale cress)
    Product Homeodomain-like superfamily protein
    Comment Comment: REVIEWED REFSEQ: This record has been curated by TAIR and Araport. This record is derived from an annotated genomic sequence (NC_003075). On Sep 12, 2016 this sequence version replaced NM_179182.1.

    1
    61
    121
    181
    241
    301
    361
    421
    481
    541
    601
    661
    721
    781
    841
    901
    961
    1021
    1081
    ATGGTTCAAA CAGAAACCGA TCAAAGGATG GGTCTTAATC TGAATTTGTC TATCTACTCT 
    TTACCAAAAC CCTTATCTCA ATTTCTCGAT GAAGTGTCGA GGATCAAAGA CAATCATTCC
    AAACTGTCTG AAATCGATGG TTATGTCGGA AAATTAGAGG AAGAAAGGAA CAAAATCGAT
    GTTTTTAAAC GAGAACTCCC TTTATGCATG CTCTTATTGA ACGAAGAGAT TGTTTTTCTC
    TGTGTAGCGA TTGGGGCGTT GAAGGATGAG GCTAGAAAAG GGTTATCACT GATGGCATCT
    AATGGGAAAT TCGATGATGT TGAAAGGGCA AAACCGGAAA CTGATAAGAA GAGCTGGATG
    AGCTCTGCCC AGTTATGGAT CTCGAACCCT AATTCCCAAT TCCGATCGAC AAATGAAGAA
    GAAGAAGATC GGTGTGTGAG TCAGAACCCA TTTCAGACAT GTAATTACCC TAATCAAGGA
    GGGGTATTTA TGCCATTTAA CCGTCCCCCA CCTCCTCCAC CACCAGCTCC TCTGTCTCTC
    ATGACTCCAA CATCAGAGAT GATGATGGAT TATAGCAGAA TTGAGCAGAG TCATCATCAT
    CATCAATTCA ATAAACCTTC TTCACAGAGT CATCACATTC AGAAGAAAGA GCAGAGAAGG
    AGATGGTCAC AGGAACTTCA CCGTAAATTT GTTGATGCTC TTCATAGGCT TGGAGGGCCA
    CAAGTTGCTA CACCAAAGCA GATTAGAGAT TTGATGAAAG TTGATGGTTT AACCAATGAT
    GAAGTCAAGA GCCATTTACA AAAATATAGA ATGCATATCC GTAAGCATCC GCTGCATCCG
    ACAAAGACTT TGTCATCGTC GGATCAGCCA GGTGTGTTAG AGAGAGAATC GCAGAGCTTA
    ATAAGTTTGT CGAGGTCGGA TTCGCCACAA AGCCCACTTG TTGCAAGAGG GTTGTTCAGT
    AGTAATGTTG GTCATAGCTC AGAGGAAGAT GAGGAGGAGG AGGATGAAGA GGAGGAGAAG
    TCTGATGGAC GTAGTAGCTG TAGAAATGAT GAAACAAAGA AGAAGAGACA AGTGTTGGAT
    CTTGAGCTTT GA

    The stop codons will be deleted if pcDNA3.1+/C-(K)DYK vector is selected.

    RefSeq NP_849513.1
    CDS85..1176
    Translation

    Target ORF information:

    RefSeq Version NM_179182.2
    Organism Arabidopsis thaliana(thale cress)
    Definition Arabidopsis thaliana Homeodomain-like superfamily protein (AT4G37180), mRNA.

    Target ORF information:

    Epitope DYKDDDDK
    Bacterial selection AMPR
    Mammalian selection NeoR
    Vector pcDNA3.1+/C-(K)DYK
    NM_179182.2

    ORF Insert Sequence:

    1
    61
    121
    181
    241
    301
    361
    421
    481
    541
    601
    661
    721
    781
    841
    901
    961
    1021
    1081
    ATGGTTCAAA CAGAAACCGA TCAAAGGATG GGTCTTAATC TGAATTTGTC TATCTACTCT 
    TTACCAAAAC CCTTATCTCA ATTTCTCGAT GAAGTGTCGA GGATCAAAGA CAATCATTCC
    AAACTGTCTG AAATCGATGG TTATGTCGGA AAATTAGAGG AAGAAAGGAA CAAAATCGAT
    GTTTTTAAAC GAGAACTCCC TTTATGCATG CTCTTATTGA ACGAAGAGAT TGTTTTTCTC
    TGTGTAGCGA TTGGGGCGTT GAAGGATGAG GCTAGAAAAG GGTTATCACT GATGGCATCT
    AATGGGAAAT TCGATGATGT TGAAAGGGCA AAACCGGAAA CTGATAAGAA GAGCTGGATG
    AGCTCTGCCC AGTTATGGAT CTCGAACCCT AATTCCCAAT TCCGATCGAC AAATGAAGAA
    GAAGAAGATC GGTGTGTGAG TCAGAACCCA TTTCAGACAT GTAATTACCC TAATCAAGGA
    GGGGTATTTA TGCCATTTAA CCGTCCCCCA CCTCCTCCAC CACCAGCTCC TCTGTCTCTC
    ATGACTCCAA CATCAGAGAT GATGATGGAT TATAGCAGAA TTGAGCAGAG TCATCATCAT
    CATCAATTCA ATAAACCTTC TTCACAGAGT CATCACATTC AGAAGAAAGA GCAGAGAAGG
    AGATGGTCAC AGGAACTTCA CCGTAAATTT GTTGATGCTC TTCATAGGCT TGGAGGGCCA
    CAAGTTGCTA CACCAAAGCA GATTAGAGAT TTGATGAAAG TTGATGGTTT AACCAATGAT
    GAAGTCAAGA GCCATTTACA AAAATATAGA ATGCATATCC GTAAGCATCC GCTGCATCCG
    ACAAAGACTT TGTCATCGTC GGATCAGCCA GGTGTGTTAG AGAGAGAATC GCAGAGCTTA
    ATAAGTTTGT CGAGGTCGGA TTCGCCACAA AGCCCACTTG TTGCAAGAGG GTTGTTCAGT
    AGTAATGTTG GTCATAGCTC AGAGGAAGAT GAGGAGGAGG AGGATGAAGA GGAGGAGAAG
    TCTGATGGAC GTAGTAGCTG TAGAAATGAT GAAACAAAGA AGAAGAGACA AGTGTTGGAT
    CTTGAGCTTT GA

    The stop codons will be deleted if pcDNA3.1+/C-(K)DYK vector is selected.

    CloneID OAb17316
    Clone ID Related Accession (Same CDS sequence) NM_119881.3
    Accession Version NM_119881.3 Latest version! Documents for ORF clone product in default vector
    Sequence Information ORF Nucleotide Sequence (Length: 1071bp)
    Protein sequence
    SNP
    Vector pcDNA3.1-C-(k)DYK or customized vector User Manual
    Clone information Clone Map MSDS
    Tag on pcDNA3.1+/C-(K)DYK C terminal DYKDDDDK tags
    ORF Insert Method CloneEZ™ Seamless cloning technology
    Insert Structure linear
    Update Date 2019-02-13
    Organism Arabidopsis thaliana(thale cress)
    Product Homeodomain-like superfamily protein
    Comment Comment: REVIEWED REFSEQ: This record has been curated by TAIR and Araport. This record is derived from an annotated genomic sequence (NC_003075). On Sep 12, 2016 this sequence version replaced NM_119881.2.

    1
    61
    121
    181
    241
    301
    361
    421
    481
    541
    601
    661
    721
    781
    841
    901
    961
    1021
    ATGGTTCAAA CAGAAACCGA TCAAAGGATG GGTCTTAATC TGAATTTGTC TATCTACTCT 
    TTACCAAAAC CCTTATCTCA ATTTCTCGAT GAAGTGTCGA GGATCAAAGA CAATCATTCC
    AAACTGTCTG AAATCGATGG TTATGTCGGA AAATTAGAGG AAGAAAGGAA CAAAATCGAT
    GTTTTTAAAC GAGAACTCCC TTTATGCATG CTCTTATTGA ACGAAGCGAT TGGGGCGTTG
    AAGGATGAGG CTAGAAAAGG GTTATCACTG ATGGCATCTA ATGGGAAATT CGATGATGTT
    GAAAGGGCAA AACCGGAAAC TGATAAGAAG AGCTGGATGA GCTCTGCCCA GTTATGGATC
    TCGAACCCTA ATTCCCAATT CCGATCGACA AATGAAGAAG AAGAAGATCG GTGTGTGAGT
    CAGAACCCAT TTCAGACATG TAATTACCCT AATCAAGGAG GGGTATTTAT GCCATTTAAC
    CGTCCCCCAC CTCCTCCACC ACCAGCTCCT CTGTCTCTCA TGACTCCAAC ATCAGAGATG
    ATGATGGATT ATAGCAGAAT TGAGCAGAGT CATCATCATC ATCAATTCAA TAAACCTTCT
    TCACAGAGTC ATCACATTCA GAAGAAAGAG CAGAGAAGGA GATGGTCACA GGAACTTCAC
    CGTAAATTTG TTGATGCTCT TCATAGGCTT GGAGGGCCAC AAGTTGCTAC ACCAAAGCAG
    ATTAGAGATT TGATGAAAGT TGATGGTTTA ACCAATGATG AAGTCAAGAG CCATTTACAA
    AAATATAGAA TGCATATCCG TAAGCATCCG CTGCATCCGA CAAAGACTTT GTCATCGTCG
    GATCAGCCAG GTGTGTTAGA GAGAGAATCG CAGAGCTTAA TAAGTTTGTC GAGGTCGGAT
    TCGCCACAAA GCCCACTTGT TGCAAGAGGG TTGTTCAGTA GTAATGTTGG TCATAGCTCA
    GAGGAAGATG AGGAGGAGGA GGATGAAGAG GAGGAGAAGT CTGATGGACG TAGTAGCTGT
    AGAAATGATG AAACAAAGAA GAAGAGACAA GTGTTGGATC TTGAGCTTTG A

    The stop codons will be deleted if pcDNA3.1+/C-(K)DYK vector is selected.

    RefSeq NP_568018.1
    CDS221..1291
    Translation

    Target ORF information:

    RefSeq Version NM_119881.3
    Organism Arabidopsis thaliana(thale cress)
    Definition Arabidopsis thaliana Homeodomain-like superfamily protein (AT4G37180), mRNA.

    Target ORF information:

    Epitope DYKDDDDK
    Bacterial selection AMPR
    Mammalian selection NeoR
    Vector pcDNA3.1+/C-(K)DYK
    NM_119881.3

    ORF Insert Sequence:

    1
    61
    121
    181
    241
    301
    361
    421
    481
    541
    601
    661
    721
    781
    841
    901
    961
    1021
    ATGGTTCAAA CAGAAACCGA TCAAAGGATG GGTCTTAATC TGAATTTGTC TATCTACTCT 
    TTACCAAAAC CCTTATCTCA ATTTCTCGAT GAAGTGTCGA GGATCAAAGA CAATCATTCC
    AAACTGTCTG AAATCGATGG TTATGTCGGA AAATTAGAGG AAGAAAGGAA CAAAATCGAT
    GTTTTTAAAC GAGAACTCCC TTTATGCATG CTCTTATTGA ACGAAGCGAT TGGGGCGTTG
    AAGGATGAGG CTAGAAAAGG GTTATCACTG ATGGCATCTA ATGGGAAATT CGATGATGTT
    GAAAGGGCAA AACCGGAAAC TGATAAGAAG AGCTGGATGA GCTCTGCCCA GTTATGGATC
    TCGAACCCTA ATTCCCAATT CCGATCGACA AATGAAGAAG AAGAAGATCG GTGTGTGAGT
    CAGAACCCAT TTCAGACATG TAATTACCCT AATCAAGGAG GGGTATTTAT GCCATTTAAC
    CGTCCCCCAC CTCCTCCACC ACCAGCTCCT CTGTCTCTCA TGACTCCAAC ATCAGAGATG
    ATGATGGATT ATAGCAGAAT TGAGCAGAGT CATCATCATC ATCAATTCAA TAAACCTTCT
    TCACAGAGTC ATCACATTCA GAAGAAAGAG CAGAGAAGGA GATGGTCACA GGAACTTCAC
    CGTAAATTTG TTGATGCTCT TCATAGGCTT GGAGGGCCAC AAGTTGCTAC ACCAAAGCAG
    ATTAGAGATT TGATGAAAGT TGATGGTTTA ACCAATGATG AAGTCAAGAG CCATTTACAA
    AAATATAGAA TGCATATCCG TAAGCATCCG CTGCATCCGA CAAAGACTTT GTCATCGTCG
    GATCAGCCAG GTGTGTTAGA GAGAGAATCG CAGAGCTTAA TAAGTTTGTC GAGGTCGGAT
    TCGCCACAAA GCCCACTTGT TGCAAGAGGG TTGTTCAGTA GTAATGTTGG TCATAGCTCA
    GAGGAAGATG AGGAGGAGGA GGATGAAGAG GAGGAGAAGT CTGATGGACG TAGTAGCTGT
    AGAAATGATG AAACAAAGAA GAAGAGACAA GTGTTGGATC TTGAGCTTTG A

    The stop codons will be deleted if pcDNA3.1+/C-(K)DYK vector is selected.

    CloneID OAb40114
    Clone ID Related Accession (Same CDS sequence) NM_001342435.1
    Accession Version NM_001342435.1 Latest version! Documents for ORF clone product in default vector
    Sequence Information ORF Nucleotide Sequence (Length: 837bp)
    Protein sequence
    SNP
    Vector pcDNA3.1-C-(k)DYK or customized vector User Manual
    Clone information Clone Map MSDS
    Tag on pcDNA3.1+/C-(K)DYK C terminal DYKDDDDK tags
    ORF Insert Method CloneEZ™ Seamless cloning technology
    Insert Structure linear
    Update Date 2019-02-13
    Organism Arabidopsis thaliana(thale cress)
    Product Homeodomain-like superfamily protein
    Comment Comment: REVIEWED REFSEQ: This record has been curated by TAIR and Araport. This record is derived from an annotated genomic sequence (NC_003075).

    1
    61
    121
    181
    241
    301
    361
    421
    481
    541
    601
    661
    721
    781
    ATGGTTCAAA CAGAAACCGA TCAAAGGATG GGTCTTAATC TGAATTTGTC TATCTACTCT 
    TTACCAAAAC CCTTATCTCA ATTTCTCGAT GAAGTGTCGA GGATCAAAGA CAATCATTCC
    AAACTGTCTG AAATCGATGG TTATGTCGGA AAATTAGAGG AAGAAAGGAA CAAAATCGAT
    GTTTTTAAAC GAGAACTCCC TTTATGCATG CTCTTATTGA ACGAAGCGAT TGGGGCGTTG
    AAGGATGAGG CTAGAAAAGG GTTATCACTG ATGGCATCTA ATGGGAAATT CGATGATGTT
    GAAAGGGCAA AACCGGAAAC TGATAAGAAG AGCTGGATGA GCTCTGCCCA GTTATGGATC
    TCGAACCCTA ATTCCCAATT CCGATCGACA AATGAAGAAG AAGAAGATCG GTGTGTGAGT
    CAGAACCCAT TTCAGACATG TAATTACCCT AATCAAGGAG GGGTATTTAT GCCATTTAAC
    CGTCCCCCAC CTCCTCCACC ACCAGCTCCT CTGTCTCTCA TGACTCCAAC ATCAGAGATG
    ATGATGGATT ATAGCAGAAT TGAGCAGAGT CATCATCATC ATCAATTCAA TAAACCTTCT
    TCACAGAGTC ATCACATTCA GAAGAAAGAG CAGAGAAGGA GATGGTCACA GGAACTTCAC
    CGTAAATTTG TTGATGCTCT TCATAGGCTT GGAGGGCCAC AAGTTGCTAC ACCAAAGCAG
    ATTAGAGATT TGATGAAAGT TGATGGTTTA ACCAATGATG AAGTCAAGAG CCATTTACAA
    GTAAAAGTTC TCATCTTTTA CAGTAACTTA AACTTTGATG TGTGGATGAA TGAATGA

    The stop codons will be deleted if pcDNA3.1+/C-(K)DYK vector is selected.

    RefSeq NP_001329349.1
    CDS108..944
    Translation

    Target ORF information:

    RefSeq Version NM_001342435.1
    Organism Arabidopsis thaliana(thale cress)
    Definition Arabidopsis thaliana Homeodomain-like superfamily protein (AT4G37180), mRNA.

    Target ORF information:

    Epitope DYKDDDDK
    Bacterial selection AMPR
    Mammalian selection NeoR
    Vector pcDNA3.1+/C-(K)DYK
    NM_001342435.1

    ORF Insert Sequence:

    1
    61
    121
    181
    241
    301
    361
    421
    481
    541
    601
    661
    721
    781
    ATGGTTCAAA CAGAAACCGA TCAAAGGATG GGTCTTAATC TGAATTTGTC TATCTACTCT 
    TTACCAAAAC CCTTATCTCA ATTTCTCGAT GAAGTGTCGA GGATCAAAGA CAATCATTCC
    AAACTGTCTG AAATCGATGG TTATGTCGGA AAATTAGAGG AAGAAAGGAA CAAAATCGAT
    GTTTTTAAAC GAGAACTCCC TTTATGCATG CTCTTATTGA ACGAAGCGAT TGGGGCGTTG
    AAGGATGAGG CTAGAAAAGG GTTATCACTG ATGGCATCTA ATGGGAAATT CGATGATGTT
    GAAAGGGCAA AACCGGAAAC TGATAAGAAG AGCTGGATGA GCTCTGCCCA GTTATGGATC
    TCGAACCCTA ATTCCCAATT CCGATCGACA AATGAAGAAG AAGAAGATCG GTGTGTGAGT
    CAGAACCCAT TTCAGACATG TAATTACCCT AATCAAGGAG GGGTATTTAT GCCATTTAAC
    CGTCCCCCAC CTCCTCCACC ACCAGCTCCT CTGTCTCTCA TGACTCCAAC ATCAGAGATG
    ATGATGGATT ATAGCAGAAT TGAGCAGAGT CATCATCATC ATCAATTCAA TAAACCTTCT
    TCACAGAGTC ATCACATTCA GAAGAAAGAG CAGAGAAGGA GATGGTCACA GGAACTTCAC
    CGTAAATTTG TTGATGCTCT TCATAGGCTT GGAGGGCCAC AAGTTGCTAC ACCAAAGCAG
    ATTAGAGATT TGATGAAAGT TGATGGTTTA ACCAATGATG AAGTCAAGAG CCATTTACAA
    GTAAAAGTTC TCATCTTTTA CAGTAACTTA AACTTTGATG TGTGGATGAA TGAATGA

    The stop codons will be deleted if pcDNA3.1+/C-(K)DYK vector is selected.

  • PubMed

    Sequence and analysis of chromosome 4 of the plant Arabidopsis thaliana.
    Nature402(6763)769-77(1999 Dec)
    Mayer K,Sch?ller C,Wambutt R,Murphy G,Volckaert G,Pohl T,D?sterh?ft A,Stiekema W,Entian KD,Terryn N,Harris B,Ansorge W,Brandt P,Grivell L,Rieger M,Weichselgartner M,de Simone V,Obermaier B,Mache R,M?ller M,Kreis M,Delseny M,Puigdomenech P,Watson M,Schmidtheini T,Reichert B,Portatelle D,Perez-Alonso M,Boutry M,Bancroft I,Vos P,Hoheisel J,Zimmermann W,Wedler H,Ridley P,Langham SA,McCullagh B,Bilham L,Robben J,Van der Schueren J,Grymonprez B,Chuang YJ,Vandenbussche F,Braeken M,Weltjens I,Voet M,Bastiaens I,Aert R,Defoor E,Weitzenegger T,Bothe G,Ramsperger U,Hilbert H,Braun M,Holzer E,Brandt A,Peters S,van Staveren M,Dirske W,Mooijman P,Klein Lankhorst R,Rose M,Hauf J,K?tter P,Berneiser S,Hempel S,Feldpausch M,Lamberth S,Van den Daele H,De Keyser A,Buysshaert C,Gielen J,Villarroel R,De Clercq R,Van Montagu M,Rogers J,Cronin A,Quail M,Bray-Allen S,Clark L,Doggett J,Hall S,Kay M,Lennard N,McLay K,Mayes R,Pettett A,Rajandream MA,Lyne M,Benes V,Rechmann S,Borkova D,Bl?cker H,Scharfe M,Grimm M,L?hnert TH,Dose S,de Haan M,Maarse A,Sch?fer M,M?ller-Auer S,Gabel C,Fuchs M,Fartmann B,Granderath K,Dauner D,Herzl A,Neumann S,Argiriou A,Vitale D,Liguori R,Piravandi E,Massenet O,Quigley F,Clabauld G,M?ndlein A,Felber R,Schnabl S,Hiller R,Schmidt W,Lecharny A,Aubourg S,Chefdor F,Cooke R,Berger C,Montfort A,Casacuberta E,Gibbons T,Weber N,Vandenbol M,Bargues M,Terol J,Torres A,Perez-Perez A,Purnelle B,Bent E,Johnson S,Tacon D,Jesse T,Heijnen L,Schwarz S,Scholler P,Heber S,Francs P,Bielke C,Frishman D,Haase D,Lemcke K,Mewes HW,Stocker S,Zaccaria P,Bevan M,Wilson RK,de la Bastide M,Habermann K,Parnell L,Dedhia N,Gnoj L,Schutz K,Huang E,Spiegel L,Sehkon M,Murray J,Sheet P,Cordes M,Abu-Threideh J,Stoneking T,Kalicki J,Graves T,Harmon G,Edwards J,Latreille P,Courtney L,Cloud J,Abbott A,Scott K,Johnson D,Minx P,Bentley D,Fulton B,Miller N,Greco T,Kemp K,Kramer J,Fulton L,Mardis E,Dante M,Pepin K,Hillier L,Nelson J,Spieth J,Ryan E,Andrews S,Geisel C,Layman D,Du H,Ali J,Berghoff A,Jones K,Drone K,Cotton M,Joshu C,Antonoiu B,Zidanic M,Strong C,Sun H,Lamar B,Yordan C,Ma P,Zhong J,Preston R,Vil D,Shekher M,Matero A,Shah R,Swaby IK,O'Shaughnessy A,Rodriguez M,Hoffmann J,Till S,Granat S,Shohdy N,Hasegawa A,Hameed A,Lodhi M,Johnson A,Chen E,Marra M,Martienssen R,McCombie WR